Scientist. Experimental Zoologist, Professor and Publisher. He developed the first successful animal-tissue cultures and pioneered organ-transplantation techniques. In a 1906 experiment, he placed a piece of a frog's embryonic nerve tissue into a drop of frog lymphatic fluid, discovered that the nerve tissue actually continued growing and did not die. For this work, he was twice considered for the Nobel Prize, but the full value of his work was not yet appreciated. Harrison was once described by Fortune as "America's most famous unknown scientist". In 1998, historians of science Meyer Friedman and Gerald W. Friedland named his discovery of a way to grow cells outside the body as one of "medicine's ten greatest discoveries." He received his BA from Johns Hopkins University in 1889 and his PhD in Zoology from the same university in 1894. At that time, he was given the Bruce fellowship in biology. According to a 1894 Baltimore Sun article, "The stipend is the income of the Adam T. Bruce endowment fund of $10,000. It is always awarded to a student who has already held an ordinary fellowship in biology." He was a teacher of Morphology at Bryn Mawr College from 1894 to 1895. He went to the University of Bonn Medical School in 1899, during the time that he was an Associate Professor of Gross Anatomy at Johns Hopkins. Yale was beginning to offer a Biological Course to students who planned to study medicine, and they lured Harrison from Hopkins by promising to make him a full professor of comparative anatomy and biology at Yale. He was a professor of anatomy at Yale from 1907 to 1938, and he also served as chairman of the zoology department. In 1913, he directed the building of the Osborn Memorial Laboratories which was at that time considered the finest facility of its type. He founded the Journal of Experimental Zoology, and served as its managing editor from 1903 to 1946. In 1913, he was elected to the National Academy of Sciences. (This is an honorific society of distinguished scholars engaged in scientific and engineering research.) He was also a member of the Royal Society which is the United Kingdom's National Academy of Science. After retiring from Yale in 1938 as Sterling Professor of Biology, he became chairman of the National Research Council. In that position he helped coordinate the efforts American and British scientists during World War II which including speeding up of the production of penicillin.